Your layout diagram indicates three fire place openings. All of them need to be checked.
Where are you taking your combustion air from?
Did the flame guide make any difference when you installed it?
ETA: You are looking for light or air currents.
There once were 3 fireplace openings on the central chimney, but one of them was sealed at the fireplace (I think it's plaster - but my husband absolutely refuses to allow me to demo it, lol). That's the one in the dining room, in my diagram. There are 2 in the library - the stove is in the beehive fireplace and then there's a small one that's sealed at the top of the firebox with sheet metal.
THEN there's another fireplace, on an exterior chimney, in the family room addition.
I need to check all of these? How do I check the one that's lined for the oil burner? (There is a sweep access door - about 8 inches square and made of cast iron - in the basement for this flue, but I've not looked into it since the chimney guys lined that flue...)
And as for combustion air, well let's just say that fresh air seems to find its way into this house easily...we've insulated the attic and tightened the windows/gaps where we see/feel them with plastic, caulk, foam and rope caulk...the front door and storm are new and well weatherstripped...but this will never be a closed air system.